Pre väčšinu ľudí sú Skipper, Kowalski, Rico a Pešiak iba štyria roztomilí tučniaci žijúci v zoologickej záhrade v Central Parku. A presne to chcú, aby si o nich ľudia mysleli. Prečo? Pretože títo na pohľad mierumilovní tučniaci sú v skutočnosti elitnou bojovou jednotkou s výsadkárskym výcvikom a tajnou centrálou.
When Santa and his reindeer crash onto the island of Madagascar it's up to Alex, Marty, Gloria, Melman and those wacky penguins to deliver the presents and save Christmas.
Bent on an agenda to take over the world, the penguins' mad dolphin nemesis, Dr. Blowhole, kidnaps King Julien in an effort to lure the flightless force into a trap so that he can annihilate them once and for all. It’s up to the penguins to spring the trap before Blowhole’s plan to flood the Earth comes to fruition.
When toy dolls are made to look like Mort at the zoo, Julien gets jealous and has them taken back. Unfortunately, this causes the actual Mort to be shipped back with the toys.
A festive "King Julien Day" is held in Juliens honor. Unfortunately, no one will come unless Maurice bribes them with candy. And as much as Skipper doesn't want to join in the party, he can't seem to avoid it.
After Mort accidentally destroys Julien's floating chair because of his obsession with his feet, the king establishes a law saying that anyone who touches his feet will be banished from the "kingdom". Mort disobeys it in his sleep, causing him to be kicked out forever. However, the penguins make him feet-phobic and he stops obsessing. Later, Julien floats away in his chair and Mort has to save him from falling into a hot dog stand. He becomes obsessed again and saves him from death.
When Julien is taken to the zoo infirmary for acting even weirder than usual, Maurice is left in charge. Everything goes smoothly, until suddenly Maurice starts acting even crazier than Julien ever did.
Rico the Penguin is known for being helpful for coughing up random items, but today a zookeeper gives him de-regurgitating medicine. Normally this wouldn't be a problem, except that Rico has a time bomb in his stomach.
At closing time, the penguins hear a little girl crying that she has lost her doll, Private goofingly thinking Skipper was crying. Skipper quickly informs Private that is not him. The penguins try to throw it to her, but this backfires when the doll hits the girl and bounces into the sewer, making the girl cry.
At the zoo during night, King Julien's boom box gets stolen. Everyone gathers up and starts to complain about their apparently stolen items. Soon, Rico talks to Private and Private tells Skipper that their television has been stolen. Skipper and the penguins go off to confront the masked thief.
Furious with King Julien skunk bombing them the baboons steal his groove, using so called "magic" so he can't dance anymore and put it in a jar. Desperate for help Julien seeks the help of the penguins to get his groove back. But when fighting over the jar that contains his groove it breaks and enters Skipper's body. The baboons will only help if King Julien apologizes. But given his stubbornness King Julien still refuses to apologize.
When a beautiful falcon crash lands in the zoo, Skipper insists that she stays to recuperate. But once she tries to eat Marlene behind their backs, Skipper and the others try to put her to a stop.
The penguins' dolphin nemesis, Dr. Blowhole, is out for revenge. After Blowhole takes King Julien hostage, Skipper and gang must rush to rescue their annoying neighbor and stop evil Dr. Blowhole before he destroys the planet.
Pre väčšinu ľudí sú Skipper, Kowalski, Rico a Pešiak iba štyria roztomilí tučniaci žijúci v zoologickej záhrade v Central Parku. A presne to chcú, aby si o nich ľudia mysleli. Prečo? Pretože títo na pohľad mierumilovní tučniaci sú v skutočnosti elitnou bojovou jednotkou s výsadkárskym výcvikom a tajnou centrálou.